When complete, the project will improve services for more than 54,000 people in Bearsden, Milngavie, Clydebank and Strathblane.
The works will involve the installation of around eight miles of new water main, the transfer of supply to Milngavie WTW and the decommissioning of Burncrooks WTW.
The main 4.3 miles section of new plastic twin-pipe water main will be installed from Bankell Service Reservoir in the east, to Baljaffray Service Reservoir in the west, following a route up part of Strathblane Road, west along the north of Craigmaddie Reservoir, round the north and west of Mugdock Reservoir, down Mugdock Road to Drumclog Car Park.
It will continue across the south of Mugdock Country Park, crossing the West Highland Way over the Allander Water near Clober, crossing the A809 just south of Craigton village, to Baljaffray Service Reservoir.
The total of eight miles of water main will also include a second stretch of 3.5 miles of 250ml pipe, to be installed from near Baljaffray to Carbeth, where it will connect to the existing network.
Scottish Water's alliance partners Caledonia Water Alliance (CWA) will carry out the work, which is expected to take around two years to complete.
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Paul Sexton, Scottish Water's alliance management general manager, said: "We are delighted to be starting this important project which will benefit more than 54,000 customers for many years to come.
"Switching supply from the ageing Burncrooks Water Treatment Works to the state-of-the-art Milngavie WTW will enable us to continue providing high-quality water to thousands of domestic properties and businesses and a large number of public buildings including more than 30 schools in three local authority areas and hospitals such as the Golden Jubilee in Clydebank."
Roseanna Cunningham, Cabinet Secretary for Environment, Climate Change and Land Reform, added: "I welcome Scottish Water's announcement of £30 million investment into the drinking water infrastructure for parts of the Bearsden, Milngavie, Clydebank and Strathblane areas, which will secure a high-quality water supply into the future.
"Importantly, the installation of new pipes will also improve the resilience of the network and security of supply so this is great news for the tens of thousands of customers in these areas."
Several other alternative routes for the pipe were considered, including through more built-up areas of Milngavie and Bearsden, but this route was selected for a number of reasons, including minimising disruption.
A new pumping station will be constructed at Bankell to pump the water along the new water mains.
Jane McKenzie, communications manager with Scottish Water, commented: "The installation of a stretch of the new water main will require the temporary closure of the Drumclog car park on Mugdock Road for a short period this summer and some temporary diversions of footpaths.
"We are exploring the possibility of alternative car parking in the surrounding area. We recognise that the temporary closure of the car park and footpaths will cause some inconvenience but would remind affected customers that our substantial investment will benefit more than 50,000 people for generations to come."
Councillor Billy Hendry, Convener of East Dunbartonshire Council's Place, Neighbourhood & Corporate Assets Committee, said: "Visitors to Mugdock Country Park should be aware that there will be some disruption to areas of the park, including Drumclog car park. This is unavoidable but temporary and we have assurances that the disruption will be minimised as far as possible.
"Scottish Water has been liaising with us through the planning and implementation stages of this development and we would like to thank our visitors in advance for their understanding as this major infrastructure project is carried out."
